An end effector for stapling patient tissue is disclosed. The end effector comprises a frame comprising a proximal end and a distal end, a plurality of staples, a firing system configured to implant the staples into the patient tissue, an implantable layer comprising a tissue-contacting surface and a support surface, and an array of securement members extending above and below the support surface of the implantable layer. The securement members are detached from the frame concurrently with the firing system being actuated to implant the staples.