A fastener cartridge can comprise a cartridge body including a plurality of cavities arranged in longitudinal rows. Some of the cavities can have fasteners removably stored therein. Other cavities can have deployable tissue engaging members stored therein which are movable from an undeployed position to a deployed position to engage tissue. In some instances, the fastener cartridge can include at least one longitudinal row of cavities which each include a deployable tissue engaging member. Such rows of cavities can comprise the outer-most rows of cavities in the fastener cartridge.