Various embodiments of a device for applying a substance to, for example, eyelashes and/or eyebrows are disclosed. The device may include a support extending along a longitudinal axis, and a first row of teeth and a second row of teeth integrally formed as a single piece with the support. Teeth in the first and second rows of teeth may extend from a common side of the support, and the first and second rows of teeth may extend over at least one fourth of a length of the support. At least two teeth may extend from the support in directions that are not parallel to one another. At least one of the first and second rows may include two teeth each extending, at least partially, on either side of a separation surface extending along that row.